Ursus arctos (Linneaus 1758 ; up to 3m in length) is the extant grizzly bear. It has a deeper face than Ursus maritimus and the two do not nest together in the LRT.
Ursavus tedfordi (Hoffman 1887, late Miocene) is a genus derived from Ursus arctos and basal to all other more derived Carnivora on this branch in the LRT.
Hofmann A 1892. Beiträe zur miocaenen Säugetierfauna der Steiermark. Jahrb Geol Reichanst, 42(1): 63−76.
Linnaeus C 1758. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.
Jiangzuo Q and Flynn JJ 2020. The earliest ursine bear demonstrates the origin of plant-dominated omnivory in Carnivora. IScience CellPress doi.org/10.1016/j.isci.2020.101235
Qiu Z-X, Deng T and Wang B-Y 2014. A late Miocene Ursavus skull from Guanghe, Gansu, China. Vertebrata PalAsiatica 52(7):265–302.
