Machaeroides, Daphoenus and Protictitherium

Daphoenus vetus (Leidy 1853; Middle Eocene to Middle Miocene, 37-16mya). Like related dogs (genus: Canis) this mid-sized predator dug burrows for offspring nesting and hiding sites.

Protictitherium crassum (Kretzoi 1938; Koufos and Konidaris 2011; Miocene-Pliocene) is traditionally considered a hyena-like civet. Civets are not related to hyenas in the LRT.. Here Protictitherium nests with Vulpavus, a basal placental.

Machaeroides eothen (Matthew 1909; Eocene, 56mya) has been difficult to nest, with some experts labeling this genus close to Oxyaena, a marsupial creodont. Here Machaeroides nests with Protictitherium and Daphoenus based on nmnhpaleobiology_338172 in the collection of the Smithsonian.

Kretzoi M 1938. Die Raubtiere von Gombaszög nebst einer Übersicht der Gesamtfauna. Annales Musei Nationalis Hungarici 31:88–157.

Leidy J 1853. Observations on a collection of fossil Mammalia and Chelonia from the Mauvaises Terres of Nebraska. Proc. Acad. Nat. Sci. Philad., 6: 392–394.

Matthew WD 1909. The Carnivora and Insectivora of the Bridger Basin, middle Eocene. Memoirs of the American Museum of Natural History 9:289-567.

wiki/Daphoenus
wiki/Protictitherium
wiki/Machaeroides