Aetobatus narinari (originally Raja narinari Euphrasén 1790; 5m in length, 3m wingspan) is the extant spotted eagle ray. Traditionally it is nested with Manta, but here nests with Squatina. That means the three tested rays are convergent.

The distinctive flat muscular snout is created by the anterior processes of the pectoral fins conjoining anteriorly. This sting ray has detachable venom spines at the base of its tail. While free swimming and capable of leaping clear of the water, the spotted eagle ray feeds on shelled invertebrates hiding beneath sea sands. Distinct from Squatina, the marginal jaws of Aetobatus are nearly toothless. The vomer and a medial plate between the dentaries include a series of flat plates acting as crushing palatal teeth distinct from other tested rays.