Tuditanus punctulatus (Cope 1871, Carroll and Baird 1968) middle Pennsylvanian ~306 mya. was derived a sister toPantylus and was basal to Utaherpeton.

Distinct from Pantylus, the jawline of Tuditanus was concave with longer teeth. The occipital notch was filled in with a posterior expansion of the squamosal and a wider skull roof that kept the original order of unfused bones as found in Utegenia. Carroll and Baird (1968) reconstructed the skull roof with a fused supratemporal/tabular, as in Pantylus, but I found these two to be unfused. It appears possible that the prefrontal and larimal were fused, as they are in Diplocaulus.

The scapulocoracoid was poorly ossified. The metacarpals were all subequal. All of the phalanges were subequal. The manual digit unguals were hour-glass shaped, not sharp.

The ilium of Tuditanus apparently has two dorsal processes but that is the result of a splitting ofthe rotated posterior process. The tarsus was better ossified. There are only two proximal tarsal elements, as in Gephyrostegus. The metatarsals were also nearly identical with each other and the phalanges. The medial centrale was absent here and smaller in the related Pantylus. The unguals were squared off according to Carroll and Baird (1988). That wasn't apparent in the photographs and no sister taxa share that character.

Llistrofus pricei (Carroll and Gaskill 1978; Bolt and Rieppel 2015; Gee et al. 2019; Early Permian) nests with Tuditanus in the LRT. The squamosal is reduced to a posterior rim, leaving a large lateral temporal fenestra. The supratemporal was misidentified as the tabular by Gee et al.

Bolt JR and Rieppel O 2015. The holotype skull of Llistrofus pricei Carroll and Gaskill, 1978 (Microsauria: Hapsidopareiontidae). Journal of Paleontology 83(3):471–483.

Carroll RL and Baird D 1968. The Carboniferous amphibian Tuditanus (Eosauravus) and the distinction between microsaurs and reptiles. American Museum novitates 2337: 1-50.

Carroll RL and Gaskill P 1978. The order Microsauria. Memoirs of the American Philosophical Society 126:1–211.

Cope ED 1871. Observations on the extinct batrachian fauna of the Carboniferous
of Linton, Ohio. Proceedings of the American Philosophical Society, 12: 177.

Gee BM, Bevitt JJ, Garbe U and Reisz 2019. New material of the ‘microsaur’ Llistrofus
from the cave deposits of Richards Spur, Oklahoma and the paleoecology of the
Hapsidopareiidae. PeerJ 7:e6327 DOI 10.7717/peerj.6327

Gregory JT, Peabody FE and Price LI 1956. Revision of the Gymnarthridae, American Permian microsaurs. Peabody Museum of Natural History (Yale University) Bulletin 10:77 pp.

Ruta M, Jeffery JE and Coates MI 2003. A supertree of early tetrapods. Proceedings of teh Royal Society, London B (2003) 270, 2507–2516 DOI 10.1098/rspb.2003.2524 online pdf

Vaughn PP 1962. 1962. The Paleozoic microsaurs as close relatives of reptiles, again. Amer. Midland Nat., vol. 67, pp. 79-84.

wiki/Tuditanus
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