Liushusaurus acanthocaudata (Evans and Wang 2010) Early Cretaceous, ~10 cm in length, was described from eight specimens and was originally and correctly nested at the base of the Scleroglossa. Derived from a sister to Calanguban, Liushusaurus nests at the base of the Iguania, includng Euposaurus and Iguana.

Overall larger and distinct from Calanguban, the skull of Liushusaurus was lower with a smaller orbit. The naris is elongte and angled lower than 30 degrees. No quadratojugal process appears on the jugal. The squamosals is a slender strip. The torso was relatively longer. The tail remained short.

The scapulocoracoid was larger and fenestrated. The hand and foot were similar in size, thus the hand was larger and more robust, and the foot was much smaller with shorter toes.

Moqisaurus pulchrum (Dong, Wang and Evans 2022, Early Cretaceous) nests with Liushusaurus in the LRT. The well-preserved pectoral girdle in the 28 new lizard provides the earliest fossil record of a mesosternal fontanelle.

Purbicella ragei (BGS GSb581, Evans et al. 2012) Late Jurassic/Early Cretaceous, ~10 cm est length. It was originally excavated more than a century ago and assigned to the genus, Paramacellodus. Evans et al. renamed it Purbicella. Their cladistic analysis nested Purbicella generally with Lacertoidea. Evans et al. based their nesting on a partial data matrix of Conrad (2008). The large reptile tree nested Purbicella with Liushusaurus.

Hongshanxi xidi (Dong, Wang, Mou, Zhang and Evans 2019; Oxfordian, earliest Late Jurassic) nests as the proximal outgroup to the Squamata, between Liushusaurus and Euposaurus + Iguana). Oddly the hands and feet of Hongshanxi are elongate, like their arboreal sisters, but the penultimate phalanges are shorter than the more proximal phalanges, distinct from their arboreal sisters. The torso is short relative to the femur length. The authors correctly note the very odd lack of a straight frontal-parietal suture. Also very odd is the open acetabulum (hip joint), which was overlooked by the authors.

Conrad JL 2008. Phylogeny and systematics of Squamata (Reptilia) based on morphology. Bulletin of the American Museum of Natural History 310:1–182.

Dong L, Wang Y, Mou L, Zhang G and Evans SW 2019. A new Jurassic lizard from China in Steyer J.-S., Augé M. L. & Métais G. (eds), Memorial Jean-Claude Rage: A life of paleo-herpetologist. Geodiversitas 41 (16): 623-641. https://doi.org/10.5252/geodiversitas2019v41a16.
http://geodiversitas.com/41/16

Dong L, Wang Y and Evans SE 2022. A new fossil lizard (Reptilia: Squamata) from
the Lower Cretaceous of eastern Inner Mongolia, China, Cretaceous Research, https://doi.org/10.1016/ j.cretres.2022.105363.

Evans SE, Jones MEH and Matsumto R 2012. A new lizard skull from the Purbeck Limestone Group (Lower Cretaceous) of England. Bull. Soc. géol. France, 2012, t. 183(6):517-524.

Evans SE and Wang Y 2010. A new lizard (Reptilia: Squamata) with exquisite preservation of soft tissue from the Lower Cretaceous of Inner Mongolia, China.

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