Lasanius.2 (undescribed; Early Silurian, size?) is derived from Birkenia, but here the mandibles are much larger. These taxa represent a second origin of jaws in vertebrates that give rise to tetrapods and humans, rather than placoderms, catfish and sharks.

Cheirolepis (holotype, Agassiz 1835; Pearson and Westoll 1979; Early to Middle Devonian, up to 55cm) nests beween Lasanius and Leptolepis in the LRT. The mandible remains gracile. The tail is strongly heterocercal. Slender pines support the lateral fins.

The other 'classic Cheirolepis' nests with Palaeonisciformes here.

Leptolepis sprattiformis (Agassiz 1843, Middle Triassic to Early Cretaceous; 6cm in length) is traditionally considered the first true bony fish. Here in the LRT, others precede it. The rostrum is short, the eyes are large and the overall size is tiny, indicating neotony and paedomorphosis. The scales lacked ganoid. Mass graves indicate they schooled. Here Leptolepis nests with Archaeomene.