Garjainia prima (Otchev 1958; PIN 2394/5-1 Early Triassic ~240 mya, 2 m. long) was derived from a sister to Fugusuchus and preceded Erythrosuchus at the base of the Erythrosuchidae. All are derived from a sister to Euparkeria in a clade here called the Euarchosauriformes.

Distinct from Fugusuchus, the skull of Garjainia had a taller rostrum that expanded anteriorly, curved over the naris and was straight thereafter. The premaxilla ascended posterior to the naris, nearly to the upper rim of the rostrum. The maxilla had a deeply convex rim armed with long, robust teeth. The postorbital length was reduced. The antorbital fenestra was smaller and diagonal in lateral view. The posterior margin of the lateral temporal fenestra was invaded by the quadratojugal and squamosal.

Distinct from Proterosuchus, the cervicals were taller and compressed. Tall spines topped the dorsal vertebrae. The torso was deeper due to longer dorsal ribs. The tail was relatively shorter.

The scapula had an hour-glass shape, the humerus was more robust. The fingers were shorter.

The ilium had a longer posterior process. The pubis and ischium were separate from each other and more ventrally oriented. All the pedal elements were shorter and composed of subequal phalanges.

Vjushkova triplocosta (von Huene 1960; Middle Triassic) was considered congeneric by Butler et al. 2019 using data shown above.

A selection of Erythrosuchids to scale. Click to enlarge.

Butler RJ, Sennikov AG, Dunne EM, Ezcurra MD, Hedrick BP, Maidment SCR, Meade LE, Raven TJ and Gower DJ 2019. Cranial anatomy and taxonomy of the erythrosuchid archosauriform ‘Vjushkovia triplicostata’ Huene, 1960, from the Early Triassic of European Russia. Royal Society Open Science 6: 191289. http://dx.doi.org/10.1098/rsos.191289

Ezcurra MD, Gower DJ, Sennikov AG and Butler RJ 2018. The osteology of the holotype of the early erythrosuchid Garjainia prima (Diapsida: Archosauromorpha) from the upper Lower Triassic of European Russia. Zoological Journal of the Linnean Society, 2018, XX, 1–67.

Otchev VG 1958. Novye dannye po pseudozukhiyam SSSR: Doklady Akademii Nauk SSR, 123(4):749-751.

Parrish JM 1992. Phylogeny of the Erythrosuchidae. Journal of Vertebrate Paleontology 12:93–102.

Huene F von 1960. Ein grosser Pseudosuchier aus der Orenurger Trias. Palaeontographica Abteilung A 114:105-111.

wiki/Garjainia
wiki/Vjushkovia