Miguashaia bureaui (Schultze 1973, Cloutier 1996; Middle Devonian; up to 45cm) was traditionally considered the sister group (outgroup) of the Actinista (coelocanths). Notably Miguashaia retains a heterocercal tail and it is usually reconstructed with lobe fins (see diagram above). The maxilla was considered taphonomically missing, but the premaxilla is wide. The dentary in the reconstruction is a standard coelacanth design, not what we see in this fossil in situ.
Here this specimen is reinterpreted as a Devonian 'flounder' with a wide skull, a tremendous underbite and an torso axially rotate 90º to lie flat of the sea floor, like a flounder. Lateral fins are missing, perhaps due to taphonomy. Phylogenetically it matches larger Middle Devonian taxa lacking post-crania. |