Tetraclaenodon puercensis(Cope 1881, Scott 1892; Paleocene) was originally considered a member of the Phenacodontidae. Here it nests basal to the flat-headed robust ungulate clade close to Agriochoerus antiquus.

Rostriamynodon grangeri (Wall and Manning 1986; AMNH 107635; Eocene) was originally considered amynodontid rhino. In the LRT it nests with oreodonts like Agriochoerus antiquus and also with Cambaytherium + Anthracobune. Note the shape of the nasals and the anterior extension of the frontals along with the wide molars and the molarized premolars.

Cambaytherium thewissi (Rose et al. 2014; Eocene, 55 mya; 45-75 lbs) was originally considered a basal perissodactyl, but nests here between Periphragnis and Rostriamynodon. The anterior palate has the same three lobes found in Hippopotamus. The posterior mandible has a long retroarticular process as in Anthracobune. Cambaytherium was found on the marine coastline of island India, close to where pakicetid aand archaeocete whales were convergently evolving from terrestrial tenrec ancestors.

Anthracobune pinfoldi (Pilgrim 1940; Kumar 1991, middle Eocene, size of a small tapir) The anthracobunids were originally considered proboscideans, like Elephas. Cooper et al. 2014 nested them with perissodactyls, like Ancodus. Here they nest with the mesonychids between Hippopotamus and the desmostylian Paleoparadoxia.

Anthracobunids have small and simple upper and lower incisors and relatively small canines. The naris opens dorsally signaling a greater aquatic niche. The retroarticular process is the largest among all mammals. The mandible is longer than the rostrum.