Cobelodus aculeatus (Zangerl and Case 1976, originally Styptobasis aculeata Cope 1891; FMNH UF576; Pennsylvanian, Middle to Late Carboniferous; 2m) is a large, shark-like taxon originally considered a holocephalian. Here it nests with Falcatus as 'the best known Paleozoic elasmobranch' (ca 1976).
Zangerl and Case note, "In the differentiation of the neurocranium and the visceral skeleton, Cobelodus presents a morphological condition not previously documented in any chondrichthyan, or, for that matter, any other fish."
The hyoid arch is not connected by ligaments in any with the mandibular arch. The authors considered this pirmitive, but reported, "Not everyone, nowadays, would agree that the aphetohyoidian condition in Cobelodus and its allies is primitive."
Large eyes suggest a deep sea niche. The pectoral fins have a 30cm trailing feeler/clasper supported by flexible cartilage. No anal fin is present. The pelvic fins are elongate. |