Bianchengichthys micros (Li et al. 2021, Latest Silurian, IVPP V27128A and B) is one of the smallest placoderms. Derived from Coccssteus, and basal to the flathead torsioned torso clade of the Devonian. Bianchengichthys has small fins and weak jaws. The oribit + naris openings are anteriorly oriented.
Bianchengichthys was reported by Li et al to be, "exquisitely preserved with mandible and
paired fins. Systematically, the new taxon
is close to the last common ancestor of bony and cartilaginous fishes."
Miguashaia bureaui (Schultze 1973, Cloutier 1996; Middle Devonian; up to 45cm) was traditionally considered the sister group (outgroup) of the Actinista (coelocanths). Notably Miguashaia retains a heterocercal tail and it is usually reconstructed with lobe fins (see diagram above). The maxilla was considered taphonomically missing, but the premaxilla is wide. The dentary in the reconstruction is a standard coelacanth design, not what we see in this fossil in situ.
Here this specimen is reinterpreted as a Devonian 'flounder' with a wide skull, a tremendous underbite and an torso axially rotate 90º to lie flat of the sea floor, like a flounder. Lateral fins are missing, perhaps due to taphonomy. Phylogenetically it matches larger Middle Devonian taxa lacking post-crania.
|