Pholidogaster pisciformes (Huxley 1862, Panchen 1975; Visean, Early Carboniferous, 340 mya) was originally considered a labyrinthodont and an anthracosaur, but here nests between Osteolepis and Colosteus among the Paratetrapoda, a clade that developed limbs independent of the Tetrapoda.
The new skull reconstruction is narrower than in Panchen 1975 to match the premaxilla and pectoral girdle. The premaxilla carried a lateral fang and the dentary had a corresponding slot for it.
The vertebral column included small bones that were basal to both dorsal fins and anal fin. The long straight unpaired bones once thought to be ribs are here identified as tall slender neural spines. The tail was little different from that found in Osteolepis, including the slight upturn, like a shark's tail.
The interclavicle and clavicles extended beneath the mandibles. No scapula or coracoid was visible. Those were tiny elements medial to the coracoid and cleithrum. The fingers did not ossify. The pelvis is well ossified with an acetabulum dorsal to the pubis. The hind limb includes metatarsals and a few digits.
The ossified scales that covered the body in Osteolepis and Colosteus are not present here.