Panderichthys rhombolepis (Gross 1941; Frasnian, Late Devonian, 380 mya; 90-130cm long). Distinct from basal taxa, like Osteolepis. Pandericthys had a wide low skull, a wide low torso, a short tail and five digits (or metacarpals). No interfrontal was present. Here it nests basal to Tiktaalik and all tetrapods.

Parmastega aelidae (Beznosov et al. 2019) was recovered from 372mya strata, making it the oldest reconstructable tetrapod. It was originally aligned with Ventastega, but here nests with Pandericthys, prior to Baphetes and Macrocephalus. A concave rostrum, elevated orbits and large size mark it as a derived taxon, despite its antiquity. The intertemporal is fused to the supratemporal. The orbit is deeper than the eyeball, which is the genesis of the confluent antorbital fenestra in related taxa.