Bavarisaurus (formerly Homoeosaurus) macrodactylus (Wagner 1852, Hoffstetter 1964, Conrad 2014, scale unknown) nests as a sister to Huehuecuetzpalli, not Homoeosaurus. It has been renamed Bavarisaurus, but is not the same genus as the Bavarisaurus gut contents of Compsognathus. It appears to have a short premaxilla and a ventrally convex mandible, but the postcrania is essentially identical to Huehuecuetzpalli.

GRS-Ref-28627 (Bolet et al. 2020; Early Cretaceous) was originally considered a second amber embalmed Oculudentavis preserving more of the torso. Here it nests basal to all Tritosauria, far from Oculudentavis. A medial crest rises from the rostrum. No antorbital fenestra is present. Distinct from other tritosaurs the clavicles are medially broad.

Bellairsia gracilis (Evans 1998; Tatanda et al 2022; Middle Jurassic, Scotland, 165mya, BMNH R12678) is another tiny protosquamate nesting with the GRS specimen in the LRT. It was revealed by µCT scanning.

Tijubina pontei (Bonfim and Marques 1997) was a tiny Early Cretaceous lizard from the Crato Formation (late Aptian) of northeast Brazil. In a recent redescription Simões (2012) reported Tijubina lacked the posteroventral and posterodorsal processes of the dentary and the tibial/fibular length equaled the femoral length. The posterior dentary teeth were robust, cylindrically based, unsculptured and bore no cuspids. Simoes (2012) nested Tijubina in a basal position among the Squamata. Reynoso (1998) reported a similar nesting for Huehuecuetzpalli. Neither considered the possibility that both specimens nested in clades outside the Squamata: the Protosquamata, outside of the Iguania + Scleroglossa, and the Tritosauria, a sister to the Rhynchocephalia.

Tijubina is distinguished by its teeth, which are larger posteriorly and shaped like cylinders instead of sharp points. Tijubina was much smaller than either Lacertulus or Huehuecuetzpalli.

Simoes (2012) described Tijubina as immature due to a imcompletely calcified joints, a wide open sternal fontanelle (hole), unfused pectoral and pelvic elements. Simoes did not realize that adult tritosaurs are likewise incompletely calcified. Unlike Huehuecuetzpalli, and despite its smaller size, the carpal elements of Tijubina were well ossified.

The pes of Tijubina had tendril-like toes, indicating an arboreal lifestyle. The tarsals were not coossified, a trait typical of many (but not all) tritosaurs.

Bolet and Evans described new material (LH 20523) they thought belonged to Scandensia, but that specimen nests with Tijubina.

Bolet A and Evans SE 2011. New material on the enigmatic Scandensia, an Early Cretaceous lizard from the Iberian Peninsula. Special Papers in Palaeontology 86:99-108.

Bonfim Júnior DC and Marques RB 1997. Um novo lagarto do Cretáceo do Brazil (Lepidosauria, Squamata, Lacertilia – Formação Santana, Aptiano da Bacia do Araripe. Anuário do Instituto do Geociencias 20:233-240

Evans SE 1998. Crown group lizards (Reptilia, Squamata) from the Middle Jurassic of the British Isles. Palaeontographica Abteilung A 250:123-154.

Bonfim-Júnior F de C and Rocha-Barbosa O 2006. A Paleoautoecologia de Tijubina pontei Bonfim-Júnior & Marques, 1997 (Lepidosauria, Squamata Basal da Formação Santana, Aptiano da Bacia do Araripe, Cretáceo Inferior do Nordeste do Brasil). Anuário do Instituto de Geociências – UFRJ ISSN 0101-9759 Vol. 29 – 2 / 2006 p. 54-65.

Simões TR 2012. Redescription of Tijubina pontei, an early cretaceous lizard (Reptilia; Squamata) from the crato formation of Brazil. An Acad Bras Cienc. Feb 2, 2012. pii: S0001-37652012005000001. [PDF].

Tatanda M, Fernandez V, Panciroli E, Evans SE and Benson RJ 2022. Synchrotron tomography of a stem lizard elucidates early squamate anatomy. Nature Oct 26 2022

Wagner A 1852. Neu-aufgefundene Saurier, Uberreste aus dem lithographischen Schiefern und dem obern Jurakalke: Abhandlungen der Bayerischen Akademieder Wissenschaften Mathematisch-naturwissenschafliche Kl, 3(6): 661-710.

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