Raeticodactylus filisurensis (Stecher 2008; BNM 14524) Upper Norian, Late Triassic, ~203 mya, wingspan 135 cm was extremely primitive, yet quite distinct from MPUM 6009 hinting at a much broader and earlier radiation of pterosaurs in the Triassic. Derived from a sister to MPUM 6009, Raeticodactylus was a sister to the Austrian specimen of Austriadactylus and the holotype of Eudimorphodon.
Distinct from MPUM 6009, the skull of Raeticodactylus was more robust and topped by a rostral crest that likely supported larger soft tissues. The naris was smaller and further back on the skull, nearly completely over the antorbital fenestra. The back of the skull sloped downward and the quadrate was deeply inclined. The jugal was much deeper, especially so below the orbit. The teeth were robust and packed tightly against one another. The mandible was deep with a ventral keel beneath the crest. The retroarticular process was extremely well-developed.
The cervicals were longer and larger.
The wing was extremely gracile, narrower than in any other pterosaur. All the elements were elongated. Thus, this pterosaur could easily stand quadrupedally while also balanced on its hind legs.
The femoral head was bent at a right angle. The hind limbs were relatively much shorter.
A partial mandible named Caviramus schesaplanensis Fröbisch and Fröbisch 2006) may be related.
Nesbitt and Hone (2010) in their misguided attempt at force-fitting pterosaurs into the archosauriforms claimed that Raeticodactylus had an antorbital fossa, a structure otherwise unknown in the Pterosauria. They did not realize that what they identified as a fossa was simply the depth dimension of the fenestra along with the dorsal skull in ventral exposure, bent over due to crushing.
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