Dorygnathus purdoni GSM 3166 (?Parapsicephalus purdoni Newton 1888, Unwin 2003) Early Jurassic ~190 mya ~21 cm skull length, was originally considered a Scaphognathus, then a Parapsicephlaus and finally a specimen of Dorygnathus. D. purdoni was derived from a sister to the R156 specimen of Dorygnathus and phylogenetically preceded Angustinaripterus.

Distinct from R156, the skull of D. purdoni was 1.5 times longer. The antorbital fenestra was twice as long as deep. The orbit was slightly smaller than the antorbital fenestra and triangular. It was constricted by a dorsally expanded jugal not much wider than tne orbit itself. The upper temporal fenestra was dorsal to the orbit and had little to no lateral exposure. The body of the pterygoid was shifted posteriorly. It had a curled lateral process and a long, straight vomer process in contact with the fused vomers. The ectopalatines had a broad medial base. The teeth are largely unknown, but alveoli indicate that the posterior teeth were robust and well spaced.

See the pterosaur family tree here.

Carpenter K, Unwin DM, Cloward K, Miles CA, and Miles C 2003. A new scaphognathine pterosaur from the Upper Jurassic Formation of Wyoming, USA. In: Buffetaut, E., and Mazin, J.- M. (eds.). Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217:45-54.

Newton ET 1888. On the skull, brain and auditory organ of a new species of pterosaurian (Scaphognathus Purdoni) from the Upper Lias near Whitby, Yorkshire. \Philosphoical Transaction of the Royal Society, London 179: 503-537.

Padian K 2009. The Early Jurassic Pterosaur Dorygnathus banthenis (Theodori, 1830) and The Early Jurassic Pterosaur Campylognathoides Strand, 1928, Special Papers in Paleontology 80, Blackwell ISBN 9781405192248

Unwin DM 2003. On the phylogeny and evolutionary history of pterosaurs. In: Buffetaut E. & J-M. Mazin, Eds. Evolution and Palaeobiology of Pterosaurs. London, Geological Society Special Publication 217: 139-190.

wiki/Dorygnathus

wiki/Harpactognathus