Orobates pabsti (Berman et al. 2004) Wolcampian, Early Permian, 1.5m in length, was derived from a sister to Limnoscelis and Tseajaia. Orobates phylogenetically preceded Diadectes and Procolophon, in agreement with Berman et al. (2004). Orobates also preceded the turtle clade beginnng with Stephanospondylus and the Lanthanosuchia beginning with Nyctiphruretus.

Distinct from Limnoscelis, the skull of Orobates was shorter overall and especially in the postorbital poertion. The naris was larger. The jugal developed a postorbital process. The teeth were not sharp, as in the Milleretta clade.

The rib cage did not flare out so widely, especially anteriorly. The anterior caudal transverse processes were elongated and turned posteriorly.

The top of the ilium had a narrow "neck" below the posterior process. The limbs were more robust.

Limnoscelis and its sisters, Solenodonsaurus and the chroniosuchids appear to have been carnivores. Turtles, like Proganochelys are omnivores as a whole with each genus preferring plants, fish, jellyfish or insects. After Lanthanosuchus, members of the Lanthanosuchidae appear to have been insectivores and trended toward a smaller overall size.

See the complete tree.

Berman, DS et al. 2004. A new diadectid (Diadectomorpha), Orobates pabsti, from the Early Permian of Central Germany. Bulletin of Carnegie Museum of Natural History 35 :1-36. doi: 10.2992/0145-9058(2004)35[1:ANDDOP]2.0.CO;2

wiki/Orobates