Nurhachius iuei (Zhou et al. 2019; Early Cretaceous) was originally considered congeneric with the holotype genus, but here nests with Pterodactylus longicollum. The rostrum is narrow, the orbit is not invaded by circumorbital processes, the teeth are lanceolate with narrow roots, convergent with istiodactylids.

Nurhachius ignaciobritoi (Wang, Kellner, Zhou & Campos 2005) IVPP V-13288, Early Cretaceous, skull length ~30 cm, ~2.5 m wingspan, was derived from a sister to Coloborhynchus, and Criorhynchus. Nurhachius was a phylogenetic predecessor to SMNK PAL 1136 and Istiodactylus.

Distinct from Criorhynchus the skull of Nurhachius further extended the rostrum and increased the size of the antorbital fenestra. If predecessors had a crest, it was greatly reduced or underdeveloped in Nurhachius. The orbit was very narrow and posteriorly slanted with a tiny sclerotic ring at the top. The upper temporal fenestra was completely above the orbit.

Distinct from Coloborhynchus, the sternal keel was very deep (if that is the keel). The humerus was shorter. Fingers I-III were smaller.

The femur was shorter. The metatarsals were robust and the pedal digits were slender, as in Zhenyuanopterus.

The ornithocheirid family tree is here. The entire pterosaur family tree is here.

The Ornithocheiridae. Click to enlarge and expand.

Wang X, Kellner AWA, Zhou Z and Campos DA 2005. Pterosaur diversity and faunal turnover in Cretaceous terrestrial ecosystems in China. Nature 437 (7060): 875–879. doi:10.1038/nature03982. PMID 16208369.

Wang X, Rodrigues T, Jiang S, Cheng X and Kellner AWA 2014. An Early Cretaceous pterosaur with an unusual mandibular crest from China and a potential novel feeding strategy. Scientific Reports 4 : 6329, pp. 1-9. | DOI: 10.1038/srep06329

Zhou X, Pegas RV, Leal MEC and Bonde N 2019. Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae. PeerJ 7:e7688 DOI 10.7717/peerj.7688

wiki/Nurhachius