Liaoningopterus gui IVPP V 13291 (Wang and Zhou 2003) Cenomanian, Early Cretaceous, ~90 mya, ~61 cm skull length was derived from a sister to Cearadactylus and was a sister to Anhanguera leaving no known descendants.

Distinct from Cearadactylus, the skull of Liaoningopterus was low anteriorly and very tall posteriorly. A very low crest surmounted the snout tip. Only one premaxillary tooth was enlarged to fang status. It is the largest tooth known for any pterosaur. The anterior dentary was expanded.

The ornithocheirid family tree is here. The entire pterosaur family tree is here.

The Ornithocheiridae. Click to enlarge and expand.

Wang X and Zhou Z 2003. Two new pterodactyloid pterosaurs from the early Cretaceous Jiufotang formation of western Liaoning, China. Vertebrata Palasiatica 41(1): 34-41.

wiki/Liaoningopterus