Eolacerta robusta (Nöth 1940) Middle Eocene, ~40 mya, was considered by Müller (2001) to be so primitive that it could not be assigned to any modern lizard family. Here Eolacerta was a sister to Tupinambis and nested with Acanthodactylus and Macrocephalosaurus.
Overall larger than and distinct from Acanthodactylus, the skull of Eolacerta had a longer rostrum and a larger upper temporal fenestra. The teeth were relatively smaller.
The neural spines were taller than each centrum. The lumbar ribs were longer.
The medially broad and fenestrated clavicles were breached medially, creating a medially forked clavicle. Manual 4.4 was longer than m4.3.
The pubis was narrower and angled anteriorly. The hind feet were relatively larger with deeper unguals.