Chlamydoselachus anguineus (Garman 1884; 2m in length) is the basalmost shark + bony fish in the LRT. On one branch sharks and sturgeons arise from this taxon. On another branch the rest of the bony fish and ultimately tetrapods arise. The anterior gill slits, that connect ventrally are retained by bony fish. The posterior five gill slits are retained by sharks.

Thrinacoselache gracia (Grogan and Lund 2008; up to 1m in length; Middle Carboniferous) nests with extant Chlamydoselachus (above) in the LRT. The skull height is 2cm. The rostrum is elongate. The fins are smaller. The skin is preserved.