Pterodactylus? [Ctenochasma] elegans? (Zittel 1882, B St 1875 XIV 501, No. 45 of Wellnhofer 1970) ~4 cm skull length, was considered a species of Pterodactylus. Here No. 45 was derived from a sister to Ctenochasma elegans and AMNH 5147. No. 45 phylogenetically preceded Gegepterus, Ctenochasma gracile and Pterodaustro.

Distinct from AMNH 5147, the skull of No. 45 was lower and upturned anteriorly. The jugal was narrower so the orbit took up a majority of the skull height. The postorbital process of the jugal was horizontal, no higher than the base of the jugal. The occiput and quadrate were likewise lowered. The cranium was larger.

The metacarpus was more robust, but the fingers were smaller. The maual unguals were small, no deeper than the phalanges and shorter than any one of them. The wing phalanges were more robust.

The pubis and ischium were separated and the pubis was narrower. The ischium was angled posteriorly. The pes was smaller than the tibia.

See more ctenochasmatids here. The dorygnathid tree is here. The pterosaur family tree is here.

Ctenochasma? elegans? (unnumbered, Lauer collection; Solnhofen Fm., Late Jurassic)

Ctenochasma? elegans? (unnumbered, Fossilienatlas.de #2042) nests with n45 in the LRT.

Tanis embryo (DePalma et al. 2019, 2022; latest Cretaceous) was originally described as an azhdarchid, but here nests with small basal ctenochasmatids from the Late Jurassic.

DePalma RA et al. (10 co-authors) 2019. A seismically induced onshore surge deposit at the KPg boundary, North Dakota. Proceedings of the National Academy of Sciences. 116(17): 8190–8199.

DePalma RA et al. (15 co-authors) 2022. A Blast from the Past: Geochemical Identity of the Chicxulub Bolide and Immediate Effects of the Impact, recorded at Tanis, North Dakota. Goddard Scientific Coloquium Spring 2022.

Wellnhofer P 1970. Die Pterodactyloidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. Abhandlungen der Bayerischen Akademie der Wissenschaften, N.F., Munich 141: 1-133.

Zittel KA 1882. Über Flugsaurier aus dem lithographischen Schiefer Bayerns. - Palaeontographica 29: 7-80.

wiki/Ctenochasma
wiki/Tanis_(fossil_site