Coloborhynchus spielbergi (Owen 1874, = Ornithocheirus clavirostris; C. spielbergi Veldmeijer 2003) RGM 401 880, Early Cretaceous was originally lumped with Ornithocheirus and much later was considered congeneric with Anhanguera by several workers (Kellner 2006). Derived from a sister to Haopterus, Coloborhynchus nests here as a phylogenetic predecessor to Criorhynchus.

Distinct from Haopterus, the skull of Coloborhynchus had an anterior crest both above the snout and below the chin. The pre-antorbital fenestra region was longer. The orbit was narrower. and raised higher, over the antorbital fenestra.

The neural spines were taller. A notarium was formed by several fused dorsals into which the scapula was articulated. The sacrals were interlocked if not fused.

The sternal complex was rather deep. The scapulocoracoid was fused. The humerus was much more gracile. The ulna and radius were also thinner, but the distal ends were expanded.

The pelvis was more robust with a more ossified ischium and a raised posterior ilium.

The ornithocheirid family tree is here. The entire pterosaur family tree is here.

The Ornithocheiridae. Click to enlarge and expand.

Ikrandraco avatar (Wang et al. 2014; Early Cretaceous, 120 mya; IVPP V18199) was originally nested between Pteranodon and Ornithocheiridae. Here it nests with Coloborhynchus deep inside the Ornithocheiridae. Pteranodon is not related.

Ikrandraco had a crest only on the mandible. The eye takes up very little of the orbit. The torso was likely no longer than the ulna.

Owen R 1874. A Monograph on the Fossil Reptilia of the Mesozoic Formations. 1. Pterosauria. The Palaeontographical Society, London. pp. 1–14 & pls 1–2.

Rodrigues T and Kellner AWA 2008. Review of the pterodactyloid pterosaur Colobrohynchus. Pp. 219–228 in: Hone, D.W.E. and Buffetaut, E. (eds), Flugsaurier: pterosaur papers in honour of Peter Wellnhofer. Zitteliana B, 28.

Veldmeijer AJ 2003. Description of Coloborhynchus spielbergi sp. nov. (Pterodactyloidea) from the Albian (Lower Cretaceous) of Brazil. Scripta Geologica, 125: 35-139.

Veldmeijer AJ, Meijer HJM and Signore M 2006. Coloborhynchus from the Lower Cretaceous Santana Formation, Brazil (Pterosauria, Pterodactyloidea, Anhangueridae); an update. www.PalArch.nl, vertebrate palaeontology, 3(2): 15-29.

Wang X-L, Rodrigues T; Jiang S-X; Cheng and Kellner AWA 2014. An Early Cretaceous pterosaur with an unusual mandibular crest from China and a potential novel feeding strategy. Scientific Reports. 4: Article number 6329.

wiki/Coloborhynchus
wiki/Ikrandraco