Fenghuangopterus lii (Lü, Fucha & Chen 2010) CYGB-0037, Middle Jurassic ~160 million years old, ~8 cm skull length, was considered an early scaphognathine, and it is very close to Scaphognathus, but it is a basal dorygnathid in the lineage of ctenochasmatids. Pterorhynchus and Scaphognathus were more distant sisters. Derived from a sister to Dorygnathus SMNS 51827 , Fenghuangopterus was a s sister to Cacibupteryx and phylogenetically preceded other putative Dorygnathus specimens.

Distinct from SMNS 51827, the skull of Fenghuangopterus had a shorter rostrum and smaller teeth, for the most part. The mandble was more gracile.

The tail was largely buried, which is why it looks so gracile here.

The scapula was shorter. The sternal complex was wider. The manus was smaller, especially the medial three digits. Manual 4.3 and m4.4 were much shorter.

The pelvis was shallower and had a substantial posterior process. The hind limbs were much longer, but the feet were not.

See the pterosaur family tree here.

The Dorygnathia is a clade that went unrecognized due to taxon exclusion in all prior studies. No one bothered to check to see if every specimen labeled Dorygnathus was distinct enough to add to a cladistic analysis. Many are. Rather than being an oddball pterosaur, best mentioned as a sidelight, Dorygnathus turns out to be the key taxon with regard to the division of the former 'Pterodactyloidea' into four separate clades. Two were derived from Dorygnathus. Another two were derived from Scaphognathus, itself a descendant of Dorygnathus via Pterorhynchus.

The Dorygnathia is defined as Dorygnathus, Pterodaustro, their last common ancestor and all of its descendants.

The two pterodactyloid-grade clades that evolved from Dorygnathus include the Ctenochasmatidae and the Azhdarchidae.

Lü J, Fucha X and Chen J 2010. A new scaphognathine pterosaur from the Middle Jurassic of western Liaoning, China. Acta Geoscientica Sinica 31(2):263–266.

wiki/Fenghuangopterus